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Boyle Heights, 1958. 12 year old Maura's femininity causes a clash between her mother Rose and her grandparents, Haim and Yetta. Across town, 12 year old Shelly quits the school play and stops eating but won't tell anyone why. Flash forward to 1966, where 21 year olds Mort and Shelly carry on a secret affair.

Maura reconciles with her lineage, both past and present; Ali talks to Leslie about her academic future; Josh looks to Buzz as a father figure; Sarah tries to tie up loose ends.

Maura makes a new friend and has a rude awakening at the music festival; Sarah experiences liberation from an unlikely source; Ali makes an unexpected journey to the past.

The Pfeffermans escape their problems on land by taking a cruise together. Though the clan spends most of the cruise apart - venturing on their own journeys of self discovery - they ultimately find themselves coming together at the Seder that almost wasn't.

A funeral brings all the Pfeffermans together and reunites them with their past - Shelley and Maura with old friends, Sarah with her ex, and Josh with his teenage babysitter - who brings a 17-year-old boy. Just when they thought there were no more secrets, the family busts out a few more.

Josh goes on a date with Rabbi Raquel while Ali explores her feminine side with a TA from Gender Studies class. The kids turn out for Maura's performance in a talent show, but her delight quickly turns to dejection.

Sarah and Tammy's big, white wedding occurs. Maura and Ali are visited by ghosts from the past. And Josh and Rabbi Raquel can't hide their big news.

In a flashback, Mort escapes with friend Mark (Bradley Whitford) to a cross-dressing retreat in the woods, and becomes captivated with a very accepting woman named Connie (Michaela Watkins). Left alone after canceling her Bat Mitzvah, 13-year-old Ali finds trouble with an older man at the beach.

Maura finally comes out to Ali, and they join Sarah on a field trip to the mall - where they all get rude lessons in womanhood. Josh is forced to confront his past thanks to questions from Syd (Carrie Brownstein) about his longtime relationship with Rita.

As they struggle to come to terms with Maura, Josh looks for answers from a rabbi named Raquel (Kathryn Hahn), and Ali looks for answers in a Gender Studies class with Syd. Sarah is left to answer questions from her own kids, who meet Maura for the first time.

Josh has a pool party for his new band, but he and his siblings are caught off-guard when Maura and Shelly arrive together; Ali reconnects with an old friend; Sarah finds her past catching up with her.

While working at an LGBT suicide hotline, Maura receives a distressing call from a young trans woman, Elizah. Afraid that Elizah will hurt herself, Maura goes on an odyssey through South Central LA to find the troubled teen and lands herself in the hospital.

Davina and Shea inspire Maura to re-imagine her past; Colton's adopted family visits, leaving Josh facing a tough decision; Ali finds herself more drawn to Leslie; Sarah tries visiting a life coach.

Relationships are tested as the Pfefferman family gather for Maura's birthday. Sarah confronts Buzzy on temple board politics and Maura makes a big announcement, experiencing pushback from both Vicki and Shelly. The night culminates in a game of Sardines, forcing tense situations into tight spaces.

The Pfefferman family gathers at Hineni to support Sarah's alternative spiritual endeavor. After the news of a close friend's untimely passing, a distressed Josh leans on Raquel for support. Ali ends the night by witnessing the pitfalls of Leslie's prejudices.

When Shelly's husband Ed goes missing, Sarah, Josh and Ali set out to find him, and instead find themselves lost in their past. Ali can no longer keep Maura's secret from Josh, which sends him into a tailspin. When Maura runs into a Pfefferman family friend she takes a stand for her new identity.

Maura finds solace with ex-wife Shelly, and the two hatch a plot to help Ed realize his wishes. Sarah spends time with her ex-husband and begins to question their breakup, while Raquel finally breaks through to Josh. And when the entire family is summoned to Shelly's, they are in for a few surprises.

On Yom Kippur, Sarah tries to make amends with Tammy; Maura tries to communicate with Davina; Josh tries to make peace with Raquel; Ali and Syd host a break-the-fast party.

Josh hits the road with an unlikely copilot, stirring up some undeniable chemistry along the way. Back at home, Shelly becomes wary of Buzzy's outlandish spending while Ali sees God during a nitrous induced trip at the dentist's office.

Josh arrives in Overland Park and delivers difficult news to Colton. Sarah oversteps her boundaries with Raquel, putting the Hineni Seder in danger. A tough conversation with Bryna brings up some big issues between Maura and Vicki.

Maura yearns to forge a new relationship but finds the connection strained. Ali is drawn to rejoin her family. Out in the vast desert Josh tries his hand at shooting a gun, leaving Shelly rattled.

Maura learns more about the trans experience; Josh heads down a reckless path; Ali and Sarah prepare for the Idyllwild Wimmin's Music Festival.

Maura and Ali uncover new truths about their family. Shelly gets bold in her improv class. Sparks fly between Sarah, Lila, and Len and they set off to hatch a plan. Josh is haunted by the past.

Maura embraces her new journey, but nothing about life as a woman is easy -- making friends, finding a new community, and above all, coming out to her children. Sarah introduces her husband (Rob Huebel) to her ex-girlfriend Tammy (Melora Hardin). Josh pays a visit to his ex-babysitter Rita, as Ali gets closer to her trainer.

Josh's bonding with Colton comes at a price; Sarah feels more like an outsider than ever following a disastrous school gala; a visit to Grandma Rose leaves Ali and Syd with questions; Maura searches for autonomy.